Eto Yoshimura

Eto (エト, Eto) is a ghoul, a member, and an executive of the Aogiri Tree and is one of the main antagonists of the supernatural horror fantasy manga/anime series Tokyo Ghoul. She is the infamous One-Eyed Owl who killed Kureo Mado's wife and Akira's mother. Her human identity was Sen Takatsuki (高槻 泉, Takatsuki Sen), a best-selling horror novelist mentioned throughout the series which both Ken Kaneki and Rize Kamishiro are fans of and read her books. In near ending of the first series it is revealed that Eto is the daughter of human named Ukina and Yoshimura, a ghoul who owns Anteiku Cafe which Ken works at. She is often seen together with a fellow Aogiri member, Tatara, as both frequently watch fights from a distance.

She was voiced by Maaya Samakoko in the Japanese dub and Lindsay Seidel in the English dub.

Early Life
She was born from the ghoul Yoshimura and human Ukina. Her mother, Ukina, was forcibly killed by Yoshimura under the command of the V organization whose Yoshimura cannot oppose. Later, Yoshimura left his infant daughter to 24th ward with Ukina's journal. She grew up in the 24th ward with other ghouls and begin to bear a great hatred to the world. This results her to cannibalize other ghouls and eventually became a kakuja. After obtaining her kakuja, she began killing the CCG investigators which led the CCG to gave her the SSS rating and the title as the infamous one-eyed owl. Yoshimura, knowing this fact, numerously covered her daughter by acting as the owl, but this got discovered by the CCG and they gave him the nickname the "Non-killing Owl", as Yoshimura only injures the CCG investigators as opposed to Eto's brutal killing.

Aogiri Arc
Eto first appears where she's sitting on top of the roof of a local CCG Compound and awaits for her comrades inside; Tatara, Noro, and the Bin Brothers. Once their all done and all of the gaurds are killed, Tatara informs Eto and the whole group leaves and head back to the Aogiri Base in the 11th Ward. Eto then attends a meeting where she sees Kaneki be brought before him by Ayato Kirishima following his attack on the Anteiku Cafe with the support of Yamori and Nico. Tatara then says that Kaneki can be used by someone else and leaves with Eto after he tells Noro that the CCG will be coming and to get prepared. Once the CCG arrives at the 11th Ward Base, Eto is seen sitting down on the roof of the base and leaves with Tatara to the bridge nearby after he assigns everyone their posts and to defend different parts of the base. During the battle, Eto sees Kaneki getting strong and after the base is destroyed, she sees Kaneki and leaves afterwards. In the Root A anime, Eto greets Kaneki when he joins the Aogiri Tree in the anime's alternate timeline.

Raid on Cochlea
Following the 11th Ward Battle, the CCG was left scattered and just started regrouping and the Aogiri Tree had launched an all-out assault on the prison. Eto participated in the attack and she helped a large division of Aogiri Soldiers break into the facilit and she slaughters all CCG Guards stationed on the upper levels of the facility. While the Aogiri Forces ravage the jail and break out all of the prisoners and give the Aogiri Tree thousands of brand new members, Eto waits on the upper floor until she decieds to come down and greet Matasaka Kamishiro and convinces him to join Aogiri as he'll meet Rize once more. She also picks up Kaneki and Ayato when escaping the faciliy after they fought against Investigators Koutaru Amon and Akira Mado in her owl form.

Raid on Kanou's Lab
Eto is one of the Aogiri Members who is seen participating in the search for the mad scientist, Akihiro Kanou. She's seen waiting while Tatara is interrogating one of Kanou's assistant nurses to see where he is at. While she waits, Eto talks to Naki about Yamori's death and how Tatara should've killed Kaneki when he was given the chance to do so. Once the information has been gathered, the group discusses what move that they should make before the CCG does and they eventually find Kanou's Laboratory.

There, Eto breaks into the facility and begins her own personal search for Kanou when she runs into a pair of Aritificial One-Eyed Ghouls, Kurona and Nashiro Yasuhisa, the personal bodyguards of Doctor Kanou who Eto just runs around them as fast as if she were teleporting. Eto then taunts the twins about their dark pasts and after she leaves them, Eto bumps into Kazuichi Banjou and proceedes to rip out some of his ribs. Finally, she meets Kanou in his control room and witnesses an intense fight between an beserk Kaneki and Amon and takes an interest in Amon and then greets Kanou and leaves with him.

Anteiku Raid
After the Laboratory Raid, Eto shows up for the first time in her human form as the famous and popular author, Sen Takatsuki. She arrives at a book signing event and after signing multiple peoples' books, she runs into Hinami Fueguchi and is befriends her for her very cute and childish nature and looks. She signs Hinami's book as a gift for Kaneki and she leaves and then chats with Hide about the possibility of writing a book about a ghoul to him. Later on, Eto as Sen runs into Hinami at a coffee shop where she's having tea with Shuu Tsukiyama and has a little chat with her while Shuu's in the bathroom and is sniffing over the rag of Kaneki's blood. Eto talks with Hinami and is able to give her some helpful advice on what to do about Kaneki and then she gives Hinami her card and walks out to the CCG's Main Office where she talks with Shinohara on the possibility of turning humans into ghouls.

On the night of the CCG's Raid in the 20th Ward. Eto and the others stay behind as they wait for the CCG to finish off Yoshimura and the rest of the Anteiku Fores there. After the CCG had finished off Yoshimura, Eto went right into the battlefield in her One-Eyed Owl form and began attacking nearby investigators. Due to the intense fight with Yoshimura, Investigators Yukinori Shinohara, Kousuke Houji, Koori Ui, and Juuzou Suzuya were all exhausted and they were soon all overrun and out-matched. Eto then stabbed Shinohara and put him into a coma and continued to beat around Juuzou who desparatly tried to take her down with no success. Finally, Squad 0 arrives and Eto is confronted by Kishou Arima. The two of them engage in a brutal fight and Squad 0's reinforcments are able to help push back the Aogiri Tree and soon Eto is forced to retreat but not before consuming Yoshimura. After the battle is over the 20th Ward lies in complete ruins, Eto regurgitates her father and breaks out of her Kakuja form and greets her father.

Post-Anteiku Raid
In the months following the Anteiku Raid, Eto is seen with both Tatara and Kanou in his laboratory where he is discussing the "loss" of Kaneki (who's really been turned into Sasaki). There, Eto glances at a container with Yoshimura in it and is then told that Kaneki will be replaced with a newer and better ghoul made from her father and abducted investigators. CCG Chairman Washuu then claims that Eto and the One-Eyed King are both the same person and must be defeated at all costs.

Nutcracker Investigation
After the time-skip, Eto remains alive and well and is still very active within the story. She appears in the Aogiri Hideout where she's with Ayato discussing how he's been hired to be a security guard for Madam A at the upcoming Auction since Aogiri gets their money from her. She also warns him that the mysterious ghoul "Floppy" could show up at the event.

The Auction
On the night of the Auction, Eto and Kanou sit outside waiting and observing as the event goes down. They wait and see the Auction go down until the CCG burst in and start open firing and killing dozens of ghouls and everyone starts to run away. Eto then sends in Seidou Takizawa, whose now been turned into an artificial one-eyed ghoul, to get rid of the CCG Troops once she's given permission to send in the owl. Once Seidou is sent in and he finds out who Sasaki is, he finds out why he was sent in.

Appearance
When she's out on missions, Eto wears a very childish attire with bandages wrapped around her body and a dark pink cloak around her body. She also wears a bright pink neckscarf with flowers on it. When she operates as her human identity of Takatsuki Sen, she wears dark blue robes around her and has a very sleepy expresion on her face. She has light skin and long green hair. Her hair is always in a very sloppy fashion but it's based off of her mother. As the One-Eyed Owl, she wears a large mask with a hole in the right side and zigzag lines down on the left. There are two lines down in near the mouth and is slanted downwards. After ten years of use, her mask has now grown horns and markings on the forehead, and has single eye right in between the eyes.

Personality
Eto has a very split personality due to her ghoul nature like many other ghouls.

Ghoul
As a ghoul she displays a sadistic personality often killing people in very bloody and brutal matters. She was willing to cut off the legs of Yukinori Shinohara in order to have him "match" woth the injuries of Juuzou Suzuya. She also taunted both Nashiro and Kurona about their parents' death and has no remorse for killing both humans and ghouls alike. This is most likely from expierences from fighting in the 24th Ward.

Human
In her human form she's the opposite. Eto has a very childish personality and is very calm and uplifting. She's also very clumsy and ditzy in her human identity often oversleeping on various occasions even druing a book signing but is happy when its prohibited by her manager. She's also very friendly especially towars Hinami where she often acts nice and kind around her and will giver her helpful advice but never sugar-coats it. She appears to be viewed as a friend by Hinami and it's possible that Eto managed to convince Hinami to join Aogiri later on as implied after the destruction of Anteiku.

Powers and Abilities

 * Ukaku Kagune: Eto's kagune is very strong as it has the ability to form feathers that can shoot bullets like her father's. She has a very diverse set of powers due to the strength of her kagune. Her kagune is very powerful and is also very diverse stemming from the acts of cannibalism that she had commited in the past.
 * Kakuja Kagune: Eto has a Kakuja Kagune as a result of committing various acts of cannabilsm and devouring many ghouls during her time in the 24th Ward. She takes the appearance of a large giant creature when she's in her owl mode and has two large blades from her shoulders. In her Kakuja form, she is able to take down dozens of CCG Troops and wip out countless opponents.
 * Advanced Kagune: Eto possess an advanced form of her kagune. In this form, she is avle to summon four monster sized claws which can rip through an entire squadron of opponents. They have a similiar shape to that of Noro's kagune is very powerful and monsterous in this form.
 * Brute Strength: Eto is the strongest ghoul so far seen in the series with a SSS Rating from the CCG. Eto has a kakuja and takes the form of a behemoth sized-armor when using it in combat.
 * Combat: Like all Aogiri Tree ghouls and Executives, Eto has powerful fighting abilities. In her owl form, she is a monster and slaughters her enemies and her kagune when in her non-owl ghoul form is also powerful as well. Her combat abilities is what made her take down ghouls in the past, allowing her to eat their kagunes and grow into a Kakuja which gives her much of her strength.

Relationships

 * Yoshimura: Eto has a very twisted relationship with her father Yoshimura. During the Anteiku Raid, she helps save her father from being killed by the CCG but she then turned him in to Kanou to be experimented on. Eto apepars to have very little, it any feelings at all for her father just as long as he provides use for the Aogiri Tree.
 * Ukina: Eto appears to not know a single thing about her mother and doesn't even know that she's dead but her diary was left by Eto's side when she was an infant so she at least knows the relationship between her parents.
 * Tatara: The exact relationship between Tatara and Eto is left unknown but they appear to have some sort've a sibling relation considering how they interact. Both of them are seen next to each other nearly all the time and always acompanies him whenever a battle breaks out. Eto seems to know a lot about Tatara however so they appear to be close to say the least.