The Fallen (Transformers)

"This planet will be dark forever."

- The Fallen after activating the Solar Harvester.

The Fallen (originally known as Megatronus Prime) appears in the Transformers franchise as the Supreme leader and founder of the Decepticons.  The Fallen is the main antagonist in Transformers: Revenge of the Fallen and the most powerful of all Transformers in the film series, despite eventually being bested by Optimus Prime. Megatron, his apprentice, serves as the secondary antagonist of the movie.

Early History
Thousands of years ago, a group of the original Transformers landed on Earth. Led by their master, Megatronus Prime, he sought to create a mighty machine called a Solar Harvester to destroy Earth's sun to use as an energon source. The other Primes had determined that such harvesters could and would not be used on planets with life. However, upon discovering sentient life on Earth, Megatronus ignored this rule and lured the other Primes to Earth. There, he stole the Matrix of Leadership and betrayed his brothers, preceding to attempt to activate the Sun Harvester alongside with group of followers. A war erupted, and a great battle was fought in the area on Earth near the Sun Harvester. In the end, the Primes prevented Megatronus from activating the Harvester and imprisoned him. From this day forward, Megatronus was referred to as "The Fallen", a name which he himself embraced. The Fallen and his corrupted followers were known as the Decepticons, while the remaining Primes became Autobots. Afterwards, the Primes sacrificed themselves to hide the Matrix in a cave made of their own bodies so that the Decepticons would never activate the Harvester.

Many years ago, Megatron came into contact with the Fallen, and he manipulated Megatron into reforming the Decepticons, and commanded him to steal the All Spark. A renewal of hostilities between the Decepticons and Autobots took place on Cybertron, which resulted in the All Spark being launched into space. The Fallen commanded Megatron to pursue it, where he crash landed on Earth. The Fallen's starship, Nemesis followed Megatron and ended up crashing on a barren world, later revealed to be one of Saturn's moons. Ultimately, the Autobots followed the Decepticons to Earth and destroyed the All Spark, killing Megatron in the process.

Transformers: Revenge of the Fallen
Two years later, Megatron was resurrected by the Constructacons and returned to the massive Decepticon starship, The Nemesis. There, Megatron met with Starscream, and then proceeding to reunite with his ancient master. The Fallen told Megatron that the All Spark was merely a vessal for the power it contained, and was now in the mind of Sam Witwicky. Megatron returned to Earth and captured Sam, but could not recieve the information he needed when Optimus Prime arrived. After a long battle, Optimus successfuly killed Grindor but was killed himself by Megatron. Feeling pleased and confident at the last Prime's death, The Fallen ordered a full scale Decepticon invasion of Earth. The Fallen himself led a group of Decepticons in the landing and the destroying of an aircraft carrier. Through the Decepticon Soundwave, The Fallen appeared on all of the earth's televised devices, demanding that the "human hive" hand Sam Witwicky over to the Decepticons, or be destroyed. This caused Sam Witwicky and his close friends to be labeled as international fugitives, and the Decepticons traveled to Egypt to prepare to activate the long lost Sun Harvester.

Instead of turning over Sam, the Autobots and the U.S. Military traveled to Egypt with the body of Optimus Prime. Sam, his friends, and a few Autobots including Mudflap, Skids, and Bumblebee are transported to Egypt by Jetfire to find the Tomb of the Primes and the Matrix of Leadership to resurrect Optimus. The U.S. military and Autobots engaged in a full scale battle outside of Cairo. To battle them, the Fallen ordered more Decepticons to land on Earth and attack them and stop Sam from getting to Optimus.

During the battle, Megatron deployed the combination of all Constructicons, Devastator to destroy the pyramid surrounding the Solar Harvester, but in the process he was destroyed by a shot from the rail gun of a nearby destroyer ship under the direction of Seymour Simmons. In the chaos of Sam and Mikaela doing their best to reach Optimus, Megatron fired a rocket at Sam's feet, knocking him unconscious. Recieving a vision of the other Primes through the matrix, the Matrix was reformed and Sam revived. Sam then revived Optimus, but the Fallen teleported to their location and stunned all of the human soldiers and Autobots nearby. He stole the Matrix from Optimus's chest and teleported into the skies.

He warped over to the Solar Harvester where Megatron awaited his master and activated it, mockingly saying that his brother Primes would never be able to stop him from his ambition. But as the countdown began, the U.S. Military acted quick to prevent the destruction of Earth. The military fired tank missiles and rocket prepelled grenades at the pyramids, which barely missed hitting The Fallen and Megatron. Using his telekinetic powers, The Fallen pulled all of the soldiers and tanks up to the top of the pyramid, then dropped them out of midair to their destruction. Seeing that he had to make the ultimate sacrifice, Jetfire offered Optimus his body parts and spark power, making him extremely powerful with flight capabilities. The Fallen pulled several rocks up to the skies to take out incoming aircraft, but Optimus dodged them, knocked Megatron off the pyramid, and tackled the Fallen. In midair, he fired a rocket which destroyed the Solar Harvester. Wrestling each other in midair, they landed in a nearby village. The Fallen commanded Optimus to join his brothers in death, and engaged Optimus. The Fallen seemed to gain the upper hand until Megatron intervened and attacked Optimus from above. Megatron's energy gun was fired at his own face, and he was sent flying through a nearby stone wall, incapacitated. The Fallen re-engaged Optimus, and realized that Optimus was more than a match for him. The Fallen managed to rip off Optimus' sword, one of his rocket thrusters and hit him in the face. Then, the Fallen pulled out his spear and attempted to stab his adversary, but Optimus grabbed it, tore out of his grib and impaled The Fallen through the shoulder with his own weapon. He then demanded the Fallen's face, ripping it off, and kicking him to the side, ripped out his spark core and crushed it, ending the apocalyptic threat. Horrified at his master's demise, Megatron swore vengeance and flew away with Starscream. With the Fallen's death, the Decepticons retreated and the Human/Autobot alliance declared victory.

All of the Fallen's dialogue in the film

 * Oh, you have much to learn my disciple, the cube, was merely a vessel, its' power, its' knowledge can never be destroyed. It can only transform. (First lines)
 * It has been absorbed by the human child. The key to saving our race now lies within his mind.
 * And you will, my apprentice, in time, for millenia, I have dreamed of returning to that wretched planet where I too was once betrayed by the ones I called my brothers.
 * Only a Prime can defeat me, and now only one remains.
 * Then the boy will lead us to him, and revenge will be ours.
 * The last prime is dead.
 * Revenge... is... mine.
 * Citizens of the human hive, your leaders have withheld the truth. You are not alone in the universe. We have lived among you, hidden, but no more. As you can see, we can destroy your cities at will, unless you turnover this boy. If you resist us, we will destroy the world as you know it.
 * My Matrix!
 * My brothers could not stop me from this. Now, I claim your sun!
 * This planet will be dark forever.
 * Die like your brothers! (last words of the movie)

Transformers: Revenge of the Fallen: The Game
The Fallen voiced by James Arnold Taylor, appears in the official game adaptation of the film Transformers: Revenge of the Fallen. He is not a playable character, but is the final boss of both campaigns, except in the Nintendo DS versions where the final boss is Optimus Prime or Megatron.

Transformers: Prime
The Fallen is briefly mentioned in the episode "Partners", when Starscream tells the Autobots that he wishes to join them. Bulkhead jokes by saying that he was "lobbing with The Fallen". In the episode "One Shall Rise", the Thirteen Primes were mentioned, as the ones who vanquished Unicron. It is implied that the Fallen was one of these. According to Ratchet, Megatron named himself after the Fallen's original name: "Megatronus".

Transformers: Exodus
Megatronus Prime is mentioned as one of the original thirteen Transformers. Megatron is shown originally calling himself Megatronus, after this particular Prime, and is a gladiator, who then takes the name Megatron after the name is shortened by cheering fans at one of his matches. Megatronus, though the namesake of the Decepticon leader Megatron in the book, holds no sway over Megatron, who took the name simply because he had none of his own and wished to be associated with power and grandeur.

Transformers: Exiles
According to the book Transformers: Exiles, Megatronus Prime commissioned Solus Prime to create the Requiem Blaster. Later, Megatronus murdered Solus, triggering the first breaking among the Thirteen.

Trivia

 * The Fallen, alongside Shockwave & Scorponok are the only three Transformers in the film series who never transformed at all, though they are portrayed with alt modes in the comic adaptations & toylines. In the toyline, The Fallen transforms into an ancient Cybertronian jet.