Ai Imari

Maria Imari, otherwise known as Ai Imari, is the main antagonist of the horror manga and anime series Pupa.

Biography
While visiting her grandfather's house at Monaki Island, Ai discovered a pair of larval monsters. For thirty years, she conducted a variety of torturous experiments on the pair of larva; soon, her grandfather created a research facility that was meant to further her research. At some point, Imari was starting to fantasize about creating a virus from the cells of the larval monsters.

During a quarrel with some of the scientists who threatened to destroy her research, the female monster escapes the facility, and impregnates Sachiko Hasegawa in order to avoid recapture. Using mimicry, the Pupa monster copies Sachiko's DNA, and reincarnates herself as a clone, dubbed Yume. As for the male monster, it eventually escaped captivity, and became Yuu. Some time after the event, Ai had a falling out with her co-researcher, Bizen, and they went separate ways.

She experimented on what was left of the larval monsters, and she created the Pupa Virus, a highly contagious disease that immediately kills its victim upon infection.

Present day
Years later, she meets Yume as she was walking home from school and warns her about the red butterflies. Upon witnessing the butterflies, Yume is transformed into a ravenous monster with a lust for human flesh. Utsutsu, believing that he could reach his sister, attempts to speak with her only tto be immediately killed. When Yume returns to her humanoid form, Ai sadistically taunts her by informing her that she had eaten her brother during her rampage. Fortunately, the virus that Yume suffered from also gave Utsutsu regenerative abilities, and he returns. She gives the two siblings a drug that temporarily represses her emergence in exchange for them getting examined. In addition, Utsutsu had to serve as his sister's source for food.

Yume enters into her Emergence form, and she goes on a rampage in order to save her brother, who was being experimented on by sadistic scientists that work for Maria. Despite the fact that Yume and Utsustu escaped thanks to Yuu's intervention, Maria manages to extract a portion of the Pupa DNA, and she impregnates herself with it, out of the sake of scientific curiosity.