The Aerodent is the boss of the Topaz Passage in Wario Land 4. It is a a small ghost-mouse (or rat) that utilizes a giant inflatable bear in order to fly high enough to drop thumbtack-shaped, spiked enemies to try and harm Wario.
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The Aerodent is the only boss who does not die when he is defeated, only falling off his balloon (which explodes behind him) upon defeat.
Wario reacts by throwing the rodent back into the inflatable bear to deflate the balloon, then attacking the bear's feet to turn it upside down and attack the mouse while he blows more air into the bear. When his health starts getting low the boss begins sending down fireballs at Wario to try and set him on fire. Upon defeat, the mouse seemingly escapes but the inflatable bear explodes behind him.
The Aerodent also appears in the twenty-eighth volume of the Super Mario-kun manga in the Wario Land 4 arc. Like in the game, the Aerodent can shoot fire at Wario to turn him into Flaming Wario. It gets heavily damaged when the Item Shopkeeper uses the Large Lips attack on it, and Wario delivers the final blow to it afterwards.
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