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King Fossa is the main antagonist of Madagascar: The Video Game, and a supporting antagonist in the Netflix original All Hail King Julien. In Madagascar: The Video Game, he serves as the final boss and playable character in the Shuffleboard minigame. He is a giant fossa who is the tyrannical leader of the Fossas. In All Hail King Julien, he is known as Bone-Fossa, a normal fossa with a bird-like skull for a crown who is the leader of the Fossa Territory.
He was voiced by Fred Tatasciore in Madagascar: The Video Game and by Kevin Michael Richardson in All Hail King Julien.
In the video game, King Fossa depicted as a muscular fossa larger than his minions with brown eyes and a white mane slanting down his back. He also stands on two legs, while his minions stand on all four, possibly because he's the leader. In the Netflix original, he's depicted as a normal fossa who looks exactly like his minions, except he has gray fur and has a skull with feathers for a crown.
Madagascar: The Video Game
When Alex walks into the Fossa territory, King Fossa and his minions show up and confront him. Alex tries to escape, but they reappear and surround him. However, the penguins appear and give him the claw ability from the power cards. After Alex defeats the fossas, King Fossa roars with frustration and begins chasing Alex. He begins to eat durian and breathe at Alex. Whevever he hits Alex, he would laugh maliciously at him. Whenever he gets hit by Alex with his mangoes, he would summon his vultures to protect him and fly with durians at him. Alex would have to keep throwing mangoes at him until his healthbar is completely empty. King Fossa once again chases Alex and traps him in the arena of doom. Whenever he gets angry, his fur becomes inflamed with rage and Alex would have to wait until he's tired out from attacking him so he can claw at him. He would summon his vultures, chase Alex and use durian breath against him whenever he attacks him. Once Alex has finished him off, Marty comes out of nowhere and lands on the fossa leader. Marty then suggests that they give him to the penguins to keep him out of trouble.
He also serves as a playable character in the Shuffleboard minigame, alongside Alex, Marty, Gloria, Melman, Wilbur, Big Louie and Crocky. The levels consist of a poolside, jungle bumper and Antarctic curve.
All Hail King Julien
In All Hail King Julien, this version is known as Bone-Fossa and is not as fierce as his video game counterpart. He only appeared in three episodes. Bone-Fossa was the leader of the Foosa Territory in Madagascar, before Mary Ann takes his place as leader throughout the majority of the series, up until the end of the series, launching a coup d'état against Mary Ann and retaking his position. He also takes advantage on situations, such as Clover's disappearance on two occasions, to launch attacks on the lemurs. He was last seen being chased by Mary Ann for his betrayal.
Although his role is small in the game, he is still the main antagonist because he is the final boss and was far more dangerous than anyone else.
King Fossa, the Zoo Hunter and Sensei Croc are the only bosses in the videogame.
King Fossa in the videogame is a slightly altered version of Diego from the Ice Age franchise.
Though Bone Fossa's role is minor in All Hail King Julien, he is the final antagonist of the show.
It is unknown what happened to Bone-Fossa after being chased by Mary Ann in the penultimate episode of the series, The End is Near. Though since the Fossa are attacking the lemurs in the first film, it is possible he is still the unseen leader of the Fossa Territory and successfully overthrew Mary Ann. If this is true, Bone-Fossa could be the unseen overarching antagonist of the first film. He may have heard of Alex the Lion "claiming territory" in the jungle and was too afraid to face him off, unlike his video-game counterpart.