|“||You destroyed everything I worked for!||„|
|~ Koto getting his revenge on King Julien.|
|“||NO!. Don't listen to her. Destroy me Babak. Right here. Right in the center mass. I'm still angling for an open casket.||„|
|~ Koto's last words|
Koto is a recurring antagonist of All Hail King Julien and the main antagonist of All Hail King Julien: Exiled.
He was voiced by Maurice LaMarche who also voiced Father.
Koto was first shown inside a barrel, fell from a waterfall and onto the snake's head. Koto then befriended him and thanked him profusely for saving his life. He then claimed to be the last of his kind because of the fossa. As Koto and King Julien stay in a cave to keep safe from a lighting storm, Clover, who was on the search for King Julien, gets into a fight with Koto as she thought he was a dangerous savage. After King Julien breaks up the fight, Koto takes care of Clover's burnt foot and says she is like the warrior queens of his tribe.
King Julien shows Koto around and continues on a date with Clover, who give each other dreamy stares. When Koto and Sage rescue the lemurs from the snake as they fight, Clover and Koto later tie the snake in a knot.
Koto returns to his kingdom where many lemurs of his kind reside. He reprimands a lemur named "Benson" for not watching after his pet snake, and he then reveals to everyone his plan to take over the kingdom.
Koto returns and brings his tough warrior subjects with him to tie up the citizens and imprison them behind a gate. As King Julien expresses his betrayed feelings, Koto tells him, because there's no way they could win. When Clover gets angry, he reveals that he's going to make her his queen.
After Pancho helps King Julien and Maurice escape, Koto takes over King Julien's plane and wears his crown. Clover and Sage look at them from a tree and Clover, as she blames herself for what happened, asks Sage for his help.
On the final episode of the fifth season, Koto and Sage ensue in a fist fight during battle. As Sage was about to finish off Koto, Clover pleads to spare him and Sage does so, until King Julien accidentally leans on Koto's statue, which causes it to fall right on Koto, killing him instantly. During the victory celebration, Koto appears as a ghost, along with the other reformed villains, and nods to King Julien before returning to the afterlife.
Koto also had a short cameo on the final episode of the series partying in the afterlife with the reformed villains during Clover and Sage's wedding.