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The Skrulls are a race of imperialistic aliens in the Marvel universe.

The Skrulls have been regularly featured in several decades of Marvel Comics, usually as antagonists of the Fantastic Four and engaged in a prolonged war with the extraterrestrial Kree. At times the race has been portrayed as a villainous one, and at other times Marvel has portrayed them as any other sentient race with its own hopes and dreams, fears and prejudices, and so on. Likewise when it comes to individuals, some portrayed as villains, others as heroes, and most in the grey zone.

The Skrulls are set to appear as the main antagonists of the 2019 superhero film Captain Marvel of the Marvel Cinematic Universe (which will be based on the Kree-Skrull War comic book storyline).

Origin

Over a billion years ago, the cosmic gods known as the Celestials came to Skrullos and experimented on the Skrulls just as they have done on millions of worlds. Three branches were created, the normal Skrulls (with the potential to produce mutants with powers and perhaps evolve into Celestials themselves), the Deviant branch (with shapeshifting abilities), and the smallest cohort of them all the Eternal branch (extreme longevity and cosmically enhanced abilities).

Eventually war raged on Skrullos between the races and it was the Deviant branch which emerged victorious exterminating the basic Skrulls and the Eternals. Only one Eternal remained, Kly'bn and he was able to persuade queen Sl'gur't (the leader of the Deviants) to become his mate and join him as the god and goddess of the Skrulls. The Deviant branch later split into two more groups, the modern Skrulls and an anomaly called the Dire Wraiths, a parasitic race that could still shapeshift and had knowledges of magic, but were not as technologically advanced as the Skrulls.

The modern Skrulls originate from the planet of Skrullos, and were originally a mercantile civilization, primarily interested in free trade and willing to share their technology with all races they deemed worthy. When they encountered a new race, they simply transformed themselves to resemble that race. The Skrull empire that resulted from these contacts was based on free trade and mutual cooperation.

The Skrulls were once a peaceful space-faring race of merchants long ago, until then then they met the Kree, a hostile culture of alien warriors. Everything changed since the unfortunate encounter with the Kree, the Skrulls decided to train themselves to combat against their sworn enemies. But now, violence and conquest are the Skrulls' way of life.

Skrull Empire

The Deviant Skrulls have established the oldest known continuous interstellar empire in the universe. They have conquered or colonized every suitable planet in the Andromeda galaxy and their empire has reached neighboring galaxies. Within this intergalactic empire are trillions, and numerous sentient species. It is an empire with a totalitarian monarchy, ruled by an emperor or empress with governors running individual worlds. Skrull society consists of castes of which the known ones are the nobility, the warrior caste, the scientist caste, the priest caste, and the worker caste.

The Skrulls have been at war with the Kree for over 10 million years. They have hunted and exterminated the diabolical, vampiric, & parasitic Dire Wraiths whenever feasible. Another long standing enemy of theirs are the Badoon, a reptilian race often allied to the Kree. They are a member of the Galactic Council, which is made of up of the main interstellar powers in the Local Group of galaxies (Shi'ar, Kree, Spartoi, Brood, & the Annihilation Wave). If there is a greater threat to the Local Group of galaxies or to the universe in general, the Skrulls have worked with the members of that Council to deal with that threat.

It appears to be the most resilient empire in the cosmos, as it has never been shown to be conquered by an outside force. It may lose planets to invaders or celestial beings, have parts of the empire devastated by invaders or the Cosmic Cube, or suffer a civil war and warlordism but never conquest of the whole empire by a foreign power. In turn their rivals, the Kree Empire was conquered twice in recent memory (1st the Shi'ar Empire and then the Phalanx) and the Shi'ar Imperium was conquered by the Kree recently.

It was the Skrulls who came closest to succeeding in conquering the Earth among all alien invaders. Neither the insectoid Brood or the techno-organic Phalanx have been shown to have had success in infiltrating the Skrull Empire, unlike the Kree & Shi'ar dominions. The Skrulls in turn had an infiltrator of theirs (Nenora) who secretly ruled for a time the Kree Empire and it was Skrull infiltrators who almost brought down the Shi'ar Imperium via atrocities they had the Shi'ar military commit on its own people. Unlike the Shi'ar (with the human mutant Vulcan) or the Kree (with the Inhuman Black Bolt), the Skrulls have not had the humiliation of a hostile foreigner taking the throne.

History

During their age-long battle with the Kree, the Skrulls discovered that Earth was a key point in the galaxy in attacking the Kree's home planet, so they were resolved to take over the earth and destroy its inhabitants. They first saw the Fantastic Four as their main threat, so they tried to dispose of them, but failed. Since then, the Skrulls have waged war on almost every hero in the Marvel universe, even launching a mass invasion by impersonating all of the main superhumans on earth.

In one attempt to get rid of the Fantastic Four the Skrulls created the Super-Skrull by taking each of the FF's DNA and putting it into one of their warriors, Kl'rt.

​Avengers: Kree-Skrull War

Kl'rt the Super-Skrull (Young Avengers Vol 1 9)

Kl'rt the Super-Skrull

The Kree Empire's Supreme Public Accuser, Ronan the Accuser had wrested control of the Kree Empire from the Supreme Intelligence and attacked the Skrulls, reigniting hostilities. The Avengers become involved once the Super Skrull kidnaps Captain Marvel, and in turn they battle a Kree Sentry robot, three of the original Skrulls that had fought the Fantastic Four, S.H.I.E.L.D. agents, rogue Inhumansand legions of Skrulls. The Supreme Intelligence ends the immediate conflict by revealing to Avengers' ally Rick Jones that he has vast mental potential, which is then used to freeze all combatants in place. The Supreme Intelligence announces that, while the Kree and Skrulls have reached genetic dead ends, the human race displays incredible untapped potential. It was revealed years later that the conflict produced a Kree-Skrull hybrid, Hulkling, who is the child of one-time lovers Captain Marvel and the Skrull Princess Anelle.

Later the cosmic being Uatu the Watcher adjudicates a duel between champions from both races: Colonel Bel-Dann of the Kree and Warlord Raksor of the Skrulls. After one year of conflict the duel remains deadlocked and is ultimately abandoned with no resolution. Years later during a conflict with Xandar that included the Fantastic Four and the Young hero Nova (Richard Rider), Emperor Dorrek VII's wife, Empress R'klll, stages a coup and kills him to become ruling Empress.

Empress R'klll's reign is short-lived, however, as the greatest blow to the Skrulls arrives in the form of the World Devourer, Galactus . After his Herald Nova (Frankie Raye) annihilates the Skrull fleet, Galactus devours Tarnax IV, killing billions of Skrulls, including R'klll and the Princess Anelle. With the destruction of the central government, the Skrull Empire has collapsed into hundreds of bickering factions. Their galaxy is filled with Governors who have declared themselves to be Emperor of the Skrulls and soon a civil war began. The Shi'ar maintain a heavy presence in the former empire as well, constantly dealing with packets of resistance.

The second Kree-Skrull War

Queen Veranke (New Avengers Vol 1 40)

Queen Veranke

Later Reed Richards discovered that yet another Governor-turned-Emperor, Zabyk, had created a genetics bomb, which, when it exploded made all the Skrulls lose their shape-shifting abilities. Whatever form they were in they were stuck in at the time of the explosion.

In desperation, since without their shapeshifting powers they were vulnerable, they managed to kidnap the herald of Galactus, Nova, in an attempt to get Galactus to destroy Kree worlds for them. Their plot failed when the Silver Surfer helped Galactus rescue Nova in exchange for his freedom from Earth.

When one of the Celestials made an appearance, this scared the Skrulls into starting a second war with the Kree to show they were not powerless. The attack was first made by Kylor, one of five governors claiming to be Emperor. He had a spy in the Kree Empire, Nenora, who gave him the early edge. They tried to hide their secret weakness at all costs from the Kree.

Kylor was eventually betrayed by Nenora, as was the entire Skrull race. She assumed power of the empire and wished to keep it for herself. It was Empress S'byll who stepped up to control the Skrulls and finish the war. She was able to resurrect the Super Skrull and he returned to find the empire in shambles. They tricked the Silver Surfer into attacking a Badoon fleet, which he was not aware were allied with the Kree. Since the Kree waged war on the Surfer, he allied himself with the Skrulls to save Zenn-La from destruction.

To restore their powers, the Super Skrulls DNA needed to be transferred to S'byll, because only a female could spread the ability to the other Skrulls. The Surfer helped them to power a machine that would return the Empress' shape shifting abilities. The device was successful. With this power she was able to restore every Skrull she touched and became the true Empress of the Skrulls.

Skrull Battleship (Young Avengers, Vol 1 12)

Skrull Battleshiip

She and Super Skrull waged war with the Kree with the help of the Silver Surfer. She replaced Super Skrull with Captain Reptyl (who much later became Reptyl Prime and he helped to change her into an emotionless reptile. But Reptyl was betrayed by one of his own and the Skrull fleet was utterly destroyed by the Kree. S'byll and Silver Surfer went alone to attack the Kree homeworld of Hala.

Gaining the knowledge that Nenora was actually a Skrull gave them the advantage. S'byll gave her shape-shifting abilities back to her, crippling her and showing the Kree her true self. A peace treaty was created.

The Skrulls eventually broke that peace treaty as they kept their military drive and prefer to fight rather than negotiate. Among the other races contained within the Skrull Empire are the Druff, Guna, Kallusina, Morani, Pheragot, Queega, Tekton, and Yirbek races. Other races remain independent, whether through treaties or open resistance. Such races include the Clegrimites, Gegku, Krylorians, Wilameanis, and Xandarians. The Skrull-Xandarian war with the Xandarians recently ended with the destruction of Xandar by the supervillainess Nebula.

Skrull Automated War Machines (X-Men v3 29)

Skrull Mecha

Skrulls are meanwhile involved in numerous other activities. They were revealed to have conquered a medieval planet and abandoned it after its inhabitants perished. Skrull Prime Ten battled the Fantastic Four and Captain Mar-Vell. The Skrulls sent their own representative to witness the fate of the , at her trial by the . There is a lone appearance of Queen Akilll of the Skrull. A lone Skrull poisoned Vera Gantor to force the Avengers to seek the Resurrection Stone on his behalf. The Skrulls sent their General Zedrao to give the dying Captain Mar-Vell the Royal Skrull Medal of Honor for having been a worthy foe of the Skrulls.

Infiltrating Earth

Some time later a Skrull called Paibok the Power-Skrull mounts a campaign to invade Earth, using another Skrull called Lyja as an infiltrator. The plan involved secretly replacing the real Alicia Masters , The Thing's blind lover. The plan fell through when she instead seduced Human Torch and the two eventually married one another. The long running facade eventually came to light and the group discovered that the real, human Alicia was actually being held captive by the Skrull nation and Lyja defects, having fallen in love with the Human Torch . The Fantastic Four rescued her, but Lyja seemingly died in the process as she sacrificed herself to save Johnny when Paibok attacked.

Paibok later revealed to Devos the Devastator that she was still alive. Paibok, with the assistance of Devos, awoke Lyja from her comatose state, also bestowing her with the power to fly and fire energy blasts from her hands (becoming "Lyja the Lazerfist"). These powers were later revealed to come from a special device implanted into her. Joining them in search of vengeance, the three arrived on Earth, and isolated and battled the Torch at Empire State University. Panicking, the Torch burst into his Nova Flame and destroyed the campus. Alongside Devos and Paibok, Lyja witnessed a battle between the Fantastic Four and an alternate Fantastic Four. However, Lyja still bore feelings for the Torch and once again betrayed her cohorts to side with the Fantastic Four which ended with the Fantastic Four destroying the "Skrull War World", the Skrulls' largest attack ship.

​The Skrulls and the X-Men

A group of Skrulls traveled to Earth where they began plotting to invade the planet once again. They were however at some point approached by Apocalypse who wanted to lure together twelve powerful mutants that he needs to ascend to godhood. At the same time, the body of the Living Monolith, who had been hurled out into the far reaches of space by Thor, is found and brought back to Earth. On Earth, a mutant Skrull named Fiz sneaked into Xavier's mansion to tell them urgent news of trouble: the Skrulls have allied themselves with someone powerful and the only thing he knows is that there are twelve mutants that can't be harmed and that something terrible is going to happen. Fiz also reveals the existence of a subtype of Skrulls that possess innate powers like Earth's mutants. Deemed outcasts by the others Skrulls, they were hunted by the Empire.

After the eventual defeat of Apocalypse, Professor Xavier left into space to train them, giving them the name Cadre K. Cadre K and Xavier returned to Earth during the Kree's attempt to overtake the planet by turning it into an intergalactic prison. A wounded Z'Cann attempted to pass on vital information to Rogue. Their physical contact affected Rogue's powers for some time. The information was received and the heroes were reunited with Bishop.

Annihilation

The Skrull Empire is the first of the major interstellar empires to be invaded by the forces of Annihilus. The Annihilation Wave's superweapon, the Harvester of Sorrow, physically destroys dozens of Skrull planets but it was eventually destroyed by Kl'rt the Super-Skrull. The Skrulls then joined Richard Rider's United Front to fight against Annihilus. In the aftermath, the Skrulls unsuccessfully attempt to convince Hulkling to become their new Emperor.

Secret Invasion

Skrull Battleship (Nova v3, 18)

Skrull Battleship

In the storyline Secret Invasion, a flailing Skrull empire, under the leadership of newcomer Queen Veranke, makes an all-out effort to infiltrate and conquer Earth. It is revealed that the Skrulls were already infiltrating Earth for quite sometime by capturing and replacing many of Earth's heroes, which was now followed by the main assault group. This wave of Skrulls forms many new Super-Skrulls , with each possessing the abilities of several different heroes. Their invasion ultimately fails, costing the life of Veranke, and reducing their numbers even further. Even their Gods are slain, and the Japanese God of Evil Amatsu-Mikaboshi takes over their Pantheon, leading to the events of Chaos War.

Infinity and a New Home

During the storyline Infinity, several planets that the Skrulls settled on began being targeted by the Builders and while the Silver Surfer tried to save most of them, the Skrulls' population was once more decimated. Kl'rt later represented the Skrull Empire as a member of the Galactic Council, allied with the Avengers in the fight against the builders. After the Avengers' victory over the Builders, Kl'rt becomes the new Emperor of the Skrulls where they settle on Tarnax II, finally finding a new homeworld after years of losses.

Traits and Abilities

Skrulls are green-skinned humanoids with large pointed ears like elves or goblins, red or green eyes, and chins with several vertical folds or wrinkles in the skin below their mouths. Skrulls are known for genetic and molecular instability, and genetic diversity, due to Celestial experimentation creating the Skrull "Deviants" (now the only surviving Skrulls). The Skrulls are known for their physical malleability and ability to change to any size, shape, or color at will, taking on the appearance but not the characteristics of other beings and objects within a volume range of .75 to 1.5 times the Skrull's original volume. Skrulls are able to assume virtually any form, be it organic (e.g., cows or wild predators ) or inorganic (e.g., lamp ).

As a result, the Skrulls excel at spying and infiltration. Skrulls are also able to use their shapeshifting abilities to form weapons (e.g., blades and clubs) with parts of their bodies, making them dangerous hand-to-hand combatants. Their sexual dimorphism is roughly the same of a human being, but their orientation is more complex since they can change their genders at will. Indeed, once casually said that, for a Skrull, changing gender is comparable to a human being changing their hair color.

Courtesy of their advanced technology, the Skrulls have also been able to augment their abilities in certain warriors, such as the Super-Skrull and War Skrulls, an elite group who with special encoding are able to emulate the powers as well as appearance of Earth's superheroes.

Alternate Realities

The Skrulls have had more success as both conquerors and in resisting conquerors in alternate realities than other alien powers, including as conquerors of Earth:

  • Earth-20604: the Skrull Empire led by the billion year old Super Skrull (whose suit of armor can replicate any powers) has conquered a million worlds and exterminated most humans via giving them the "Skrull pill" which gave them super powers but eventually killed them.
  • Earth-14118: the united Skrull & Kree empire, ruled by the Skrull-Kree hybrid Dorrek VIII (son of Skrull princess Anelle & Kree warrior Mar-Vell) exterminates all life on Earth in the mid-21st century.
  • Earth-32098: in the future, the Skrulls will be allies of the X-Men. Professor Xavier establishes an intergalactic alliance of mutants (Universal X-Alliance) with the X-Men and the Skrull Cadre K forming its core.
  • Earth-32000: in the near future, the Skrulls will no longer be hostile toward Earth and become allies of the X-Men. They come to the aid of the X-Men when Earth is threatened by Apocalypse and his Shi'ar allies.
  • Earth-3290: the Earth (and its superhuman defenders) surrendered to the Skrulls at the end of Queen Veranke's Secret Invasion.
  • Earth-5692: the Skrulls conquered the Earth before the emergence of super heroes and abandoned it during the 20th century at the approach of Galactus.
  • Earth-10021: the Skrulls conquered Earth during the Secret Invasion, convert billions of humans into Skrull-Human hybrids, and those billions die later due to a virus released by Norman Osborn.
  • Earth-10219: the Skrulls' Secret Invasion stayed secret and the Skrulls are steadily replacing super heroes, leaders, etc and manipulating human affairs from the shadows.
  • Earth-804: the Skrull-Kree War is ended by the Rick Jones (actually the Kree Supreme Intelligence), Captain Mar-Vell is made the new ruler of the Kree, princess Anelle becomes the Empress of the Skrulls, and peace is made between the two.
  • Earth-9812: The Skrulls unlike the Kree and Shi'ar were not shown to have been conquered by the intergalactic Terran Empire with its Avengers Corps (Galactic Avengers Battalions).
  • Earth-6311: The Skrulls unlike the Badoon, Shi'ar, and the Universal Church of the Truth were not shown to have been conquered by Kang the Conqueror (his home reality).
  • Earth-93165: the Skrulls along with the other interstellar powers were conquered by the Kree who establish their supremacy via the Omni-Wave Projector (which destroyed the Earth). Later a coalition of rebels from various alien races and surviving Avengers overthrew the fascist Kree.
  • Earth-906: a cancerous plague unknowingly spread by the Kree Captain Mar-Vell devastated the known universe, weakening the Kree and Skrull empires and thereby putting an end to the Kree-Skrull War.
  • Earth-90111: the fascist Terran empire of the sentient android Vision conquers the Skrulls, Kree, Shi'ar, Badoon, Rigellians, and Aakon.
  • Earth-91955: In this universe which was conquered by Thanos (with the Infinity Gauntlet), the Skrulls managed to resist him for 2 hours unlike the Kree who fell within seconds.

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Bestman Salvage
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The Grandmaster | Topaz | Sakaaran Mercenaries

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Surtur | Fire Dragon

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Sambisan Captain

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Doctor Strange's variants: Strange Supreme | Sinister Strange
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Phase 5
Lord Krylar | Quantumnauts | Killer Snowbot | Vickie Tyson

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           Fantastic Four logoVillains

Comics
Abomination | Abraxas | A.I.M. | Air-Walker | Annihilation Wave | Annihilus | Apocalypse | Arcade | Ares | Attuma | Awesome Android | Baron Mordo | Basilisk | Belasco | Beetle | Beyonder | Bi-Beast | Black Knight | Blastaar | Black Cat | Brood | Brute | Bulldozer | Bullseye | Captain Barracuda | Celestials | Collector | Constrictor | Count Nefaria | Crimson Dynamo | Dark Avengers | Destroyer | Devos the Devastator | Diablo | Doctor Doom | Doctor Faustus | Doctor Octopus | Doctor Sun | Dormammu | Dragon Man | Ego | Electro | Elements of Doom | Emma Frost | Enchantress | Enclave | Fin Fang Foom | Firelord | Fixer | Frightful Four | Fu Manchu | Galactus | Giganto | Glenn Talbot | Grandmaster | Graviton | Green Goblin | Grey Gargoyle | Gorgon | Hammerhead | Harry Osborn | Hate-Monger | Hera | High Evolutionary | Hobgolin | Horsemen of Apocalypse | Hydro-Man | Hyperstorm | Immortus | Impossible Man | Ironclad | J. Jonah Jameson | Kangaroo | Kang the Conqueror | Kingpin | Klaw | Knull | Kraven | Kree | Kristoff Vernard | Living Laser | Lizard | Llyra | Loki Laufeyson | Lucia Von Bardas | Mac Gargan | Machinesmith | Mad Thinker | Maestro | Magneto | Magus | The Maker | Man-Ape | Man-Wolf | Mandrill | Mandarin | Master Pandemonium | Masters of Evil | Maximus | Medusa | Mephisto | Melter | Miracle Man | M.O.D.O.K. | Modulus | Molecule Man | Mole Man | Molten Man-Thing | Moonstone | Morgan Le Fay | Mysterio | Namor McKenzie | Nebula | Nightmare | Norman Osborn | Occulus | Onslaught | Overmind | Owl | Paibok | Piledriver | Porcupine | Power Broker | Psycho-Man | Punisher | Punisher (Earth-95126 | Puppet Master | Quasimodo | Quicksilver | Red Ghost | Red Skull | Rhino | Rogue | Ronan the Accuser | Sabretooth | Salamandra | Salem's Seven | Sandman | Sauron | Scarlet Witch | Sentinels | Shocker | Silver Sable | Sinister Six | Skrulls | Skurge | Street | Stepford Cuckoos | Super-Adaptoid | Super-Apes | Super-Skrull | Supreme Intelligence | Symbiotes | Taskmaster | Talos the Untamed | Terminus | Terrax the Tamer | Thanos | Thundra | Thunderball | Thunderbolt Ross | Thunderbolts | Tiger Shark | Tinkerer | Titania | Trapster | Tyrannus | Unicorn | Ultron | Venom | Vermin | Vulture | Warlord Krang | Whirlwind | Wizard | Wrecker | Yellow Claw | Zarrko | Zombie Mr. Fantastic

Movies
Fantastic Four (1994): Doctor Doom | The Jeweler
Fantastic Four (2005): Victor von Doom | Ned Cecil
Fantastic Four: Rise of the Silver Surfer: Galactus | Silver Surfer | Victor von Doom
Fantastic Four (2015): Victor Von Doom | Harvey Allen

Television
Fantastic Four (1967): Doctor Doom | Mole Man | Diablo | Giganto | Galactus | Blastaar | Rama-Tut | Attuma | Prince Triton | Skrulls (Super-Skrull) | Klaw | Red Ghost | Super-Apes | Molecule Man
The New Fantastic Four: Doctor Doom | Magneto | Skrulls | Mole Man | Frightful Four (Wizard, Medusa, Trapster & Sandman) | Blastaar | Impossible Man
Fantastic Four (1994): Doctor Doom | Galactus | Mole Man | Puppet Master | Warlord Krang | Super-Skrull | Frightful Four (Wizard, Trapster, Medusa & Hydro-Man) | Impossible Man | Ego the Living Planet | Maximus | Blastaar | Psycho-Man | Terrax the Tamer | Namor | Klaw | Skrulls | Annihilus
Fantastic Four: World's Greatest Heroes: Victor von Doom | Mole Man | Puppet Master | Super-Skrull | Skrulls | Supreme Intelligence | Frightful Four (Wizard & Trapster) | Attuma | Namor | Giganto | Impossible Man | Lucia Von Bardas | Terminus | Klaw | Diablo | Grandmaster | Thunderbolt Ross | Ronan the Accuser | Kree | Annihilus

Video Games
Fantastic Four (1997): Mole Man | Giganto | Psycho-Man | Skrulls (Super-Skrull) | Attuma | Namor | Doctor Doom | Galactus
Fantastic Four (2005): Victor von Doom | Annihilus | Blastaar | Puppet Master | Diablo | Mole Man
Fantastic Four: Flame On: Galactus | Skrulls
Fantastic Four: Rise of the Silver Surfer: Galactus | Victor von Doom | Terrax the Tamer | Red Ghost | Super-Apes | Super-Skrull

See Also
Future Foundation Villains | Namor the Sub-Mariner Villains | Silver Surfer Villains

           Avengers Villains

Comics
Abomination | Abominatrix | Absorbing Man | Adolf Hitler | Agony | A.I.M. | Air-Walker | Alkhema | Aleksander Lukin | Alistair Smythe | Amatsu-Mikaboshi | Amora | Andrew Forson | Anaconda | Annihilus | Ani-Men | Answer | Apocalypse | Arcade | Ares | Arnim Zola | Atlas | Attuma | Ashen Combine | Avalanche | Awesome Android | Badoon | Barracuda | Baron Blood | Baron Mordo | Baron Wolfgang von Strucker | Baron Zemo | Basilisk | Batroc | Beetle | Beyonder | Bi-Beast | Black Knight | Blackout | Black Order | Black Cat | Black Mamba | Black Talon | Black Widow | Blackie Drago | Blastaar | Blitzkrieg | Blizzard | Blob | Blood Brothers | Bombshell | Boomerang | Brothers Grimm | Brotherhood of Evil Mutants | Bulldozer | Bullseye | Burglar | Bushmaster | Bushwacker | Cache | Cannibals (Marvel Universe Vs) | Carnage | Carrion | Calypso | Celestials | Centurion | Chameleon | Chance | Chemistro | Chester Goudal | Chitauri | Collector | Colonel Ross Whittaker | Commander Kraken | Constrictor | Corruptor | Cottonmouth | Count Nefaria | Cowled Commander | Crime-Master | Crimson Dynamo | Crimson Widow | Cristu Bulat | Crossbones | Crossfire | D'Spayre | Daken | Damage | Daniel Whitehall | Dark Avengers | Dark Elves | Dark Phoenix | Deadpool | Deathbird | Deathurge | Death Adder | Destroyer | Deke Wainscroft | Destiny | Demogoblin | Diablo | Diamondback | Doctor Doom | Doctor Faustus | Doctor Octopus | Doctor Spectrum | Dormammu | Dracula | Dreadknight | Druig | Eddie Brock | Eel | Egghead | Ego the Living Planet | Electro | Elementals | Elements of Doom | Emma Frost | Enclave | Enforcers | Equinox | Erik Killmonger | Exodus | Ezekiel Stane | Fabian Cortez | Famine | Fin Fang Foom | Finn Cooley | Firelord | Fixer | Fold | Frost | Frost Giants | Frightful Four | Galactus | Gargantus | Garthan Saal | Ghost | Giganto | Goliath | Gladiator | Godzilla | Grand Director | Grandmaster | Grant Ward | Graviton | Green Goblin (Norman Osborn, Harry Osborn) | Grey Gargoyle | Grey Goblin | Griffin | Grim Reaper | Grizzly | Growing Man | Grotesk | Hammerhead | Hand | Hazmat | Hela | Hera | Herr Kleiser | Heinz Kruger | High Evolutionary | Hitman | Hive | Horsemen of Apocalypse | Hood | Hydro-Man | HYDRA | Immortus | Impossible Man | Inner Demons | Ironclad | Iron Monger | Jackal | J. Jonah Jameson | Jack O' Lantern | Jester | Jigsaw | Johnny Ohm | Jonas Hambleton | Jormungandr | Juggernaut | Justin Hammer | Kaine | Kang the Conqueror | Killer Shrike | King Cobra | Kingpin | Klaw | Knull | Korath the Pursuer | Korvac | Kraven the Hunter | Kree | Kristoff Vernard | Lady Deathstrike | The Leader | Lightmaster | Living Brain | Living Laser | Living Monolith | Loki Laufeyson | Lucia Von Bardas | Ma Gnucci | Machinesmith | Mad Pharaoh | Mad Thinker | Madame Masque | Madame Viper | Madcap | Madelyne Pryor | Maelstrom | Maestro | Maginty | Magneto | Magus | The Maker | Malekith the Accursed | Man-Ape | Mandarin | Mandrill | Man-Spider | Man-Wolf | Master Man | Master Pandemonium | Masters of Evil | Masked Marauder | Maximus the Mad | Medusa | Melter | Menace | Mentallo | Mercurio the 4-D Man | Mesmero | Mephisto | Mister Fear | Mister Hyde | Mister Negative | Mister Payback | Mister Sinister | M.O.D.A.M. | M.O.D.O.K. | M.O.D.O.K. Superior | Mojo | Molecule Man | Mole Man | Molten Man | Monica Rappaccinni | Moonstone | Morbius the Living Vampire | Morgan le Fay | Moses Magnum | Mystique | Mysterio | Namor | Nebula | Nekra | Nicky Cavella | Nightmare | Nightshade | Niles Van Roekel | Nicky Cavella | Omega Red | Onslaught | Orchis | Orka | Overdrive | Owl | Paladin | Phalanx | Phil Urich | Piledriver | Pluto | Presence | Princess Python | Proctor | Psycho-Man | Punisher | Puppet Master | Puma | Purple Man | Pyro | Quasimodo | Quicksand | Quicksilver | Radioactive Man | Ragnarok | Ramrod | Rattler | Reavers | Red Ghost | Red Skull | Reverend Samuel Smith | Rhino | Rhino II | The Rose | Ringer | Ringmaster | Riot | Rogue | Ronan the Accuser | Roxxon | Sabretooth | Sandman | Säurespritze | Sauron | Savage Land Mutates | Scarecrow | Scarlet Witch | Scorcher | Scorpion | Screaming Mimi | Sebastian Shaw | Secret Empire | Selene Gallio | Sentinels | Sentry 459 | Serpent Society | Shriek | Shocker | Shockwave | Sidewinder | Silver Sable | Silver Samurai | Sin | Sinister Six | Skaar | Skeleton Crew | Skrulls | Skurge | Slug | Space Phantom | Spider-Man Revenge Squad | Spider-Slayers | Spot | Squadron Sinister | Stranger | Street | Sunstroke | Superia | Super-Adaptoid | Super-Apes | Super Patriot | Super-Skrull | Supreme Intelligence | Superia | Surtur | Swarm | Swordsman | Symbiotes | Tarantula | Taskmaster | Technovore | Temugin | Ten Rings | Terminus | Thanos | Thorr Odinson | Thunderball | Thunderbolts | Thunderbolt Ross | Thundersword | Tiger Shark | Tinkerer | Titania | Titanium Man | Toad | Tombstone | Trapster | Trick Shot | Terrax the Tamer | Typhoid Mary | Tyrannus | U-Foes | Ulik | Ultimus | Ultron | Unicorn | Universal Church of Truth | Uranos | Vapor | Vector | Venom | Vermin | Vulture | Walrus | Warlord Krang | Whiplash | Whirlwind | White Rabbit | Will-O'-The-Wisp | Winter Soldier | Wizard | Wonder Man | Wrecker | Worthy | X-Ray | Yellow Claw | Ymir | Yon-Rogg | Zahnmörder | Zodiac | Zombie Giant-Man | Zombie Sentry | Zzzax

Movies
Marvel Animated Features
Ultimate Avengers 1 & 2: Herr Kleiser | Chitauri
Next Avengers: Heroes of Tomorrow: Ultron

Marvel Cinematic Universe
The Avengers: Loki Laufeyson | Chitauri & Leviathans | HYDRA (Gideon Malick & Jasper Sitwell) | Georgi Luchkov | Thanos
Avengers: Age of Ultron: Ultron | Ultron Sentinels | HYDRA (Wolfgang von Strucker, List, Scarlet Witch & Quicksilver) | Ulysses Klaue | Madame B | Chitauri (Leviathans) | Thanos
Avengers: Infinity War: Black Order (Thanos, Ebony Maw, Proxima Midnight, Corvus Glaive, Cull Obsidian, Nebula, Outriders, Chitauri & Leviathans) | Red Skull | The Collector | Loki Laufeyson | Winter Soldier | M'Baku | Scarlet Witch | Thunderbolt Ross
Avengers: Endgame: Black Order (Thanos, Ebony Maw, Proxima Midnight, Corvus Glaive, Cull Obsidian, Nebula, Outriders, Sakaaran Mercenaries, Chitauri, Leviathans & Chitauri Gorillas) | HYDRA/STRIKE (Red Skull, Alexander Pierce, Crossbones, Jasper Sitwell & Jack Rollins) | Loki Laufeyson (Variant L1130) | Winter Soldier | M'Baku | Scarlet Witch | Kraglin Obfonteri | Dark Elves | Thunderbolt Ross | Akihiko

Television
The Avengers: United They Stand
Ultron | Kang the Conqueror | Swordsman | Ringmaster | Masters of Evil (Baron Zemo, Absorbing Man, Boomerang, Moonstone, Tiger Shark & Whirlwind) | Attuma | Zodiac (Taurus & Scorpio) | Egghead | Scarlet Witch | Grim Reaper | Quicksilver | Namor | Doctor Doom | Baron Strucker | HYDRA | A.I.M. | Black Knight | The Collector

The Super Hero Squad Show
Doctor Doom | Mole Man | Fin Fang Foom | Abomination | Wrecker | Thunderball | Piledriver | Bulldozer | M.O.D.O.K | Klaw | Toad | Melter | Dormammu | Sabretooth | Juggernaut | Pyro | Whirlwind | Zzzax | Enchantress | Baron Mordo | Punisher | Egghead | Ringmaster | Skrulls (Super-Skrull) | Trapster | Mystique | Impossible Man | The Leader | Mad Thinker | Skurge | Crimson Dynamo | Batroc the Leaper | Loki Laufeyson | Magneto | Quicksilver | Scarlet Witch | HYDRA (Red Skull , Arnim Zola & Baron Wolfgang von Strucker) | Galactus | Terrax the Tamer | Firelord | Stardust | Thanos | Kree | Ego the Living Planet | Grandmaster | Annihilus | Nebula | Ronan the Accuser | Nightmare | Ultron | Sentinels | Dracula | High Evolutionary | Screaming Mimi

Avengers: Earth's Mightiest Heroes
HYDRA (Red Skull, Baron Von Strucker, Grim Reaper, Arnim Zola, Doughboy & Madame Viper) | Masters of Evil (Baron Zemo, Enchantress, Skurge, Chemistro, Abomination, Crimson Dynamo, Wonder Man, Living Laser & Grey Gargoyle) | A.I.M. (M.O.D.O.K., Scientist Supreme) | Kree Empire (Supreme Intelligence, Colonel Yon-Rogg & Ronan the Accuser) | Absorbing Man | Klaw | Ultron | Wrecker | Thunderball | Piledriver | Bulldozer | Red Hulk | Frost Giants (Loki Laufeyson & Ymir) | Technovore | Bi-Beast | The Leader | Whirlwind | Kang the Conqueror | Man-Ape | Malice | Mad Thinker | Mandrill | Graviton | Fenris Wolf | Lucia Von Bardas | Zzzax | Blizzard | Constrictor | Griffin | Living Laser | Whiplash | King Cobra | Magneto | Wendigo | Skrulls (Queen Veranke, Captain America Skrull, Super-Skrull) | Ulik | Malekith the Accursed | Hela | Doctor Doom | Surtur | Glenn Talbot | Winter Soldier | J. Jonah Jameson | Destroyer | Annihilus | Purple Man | Galactus | Firelord | Stardust | Terrax the Tamer | Red Ghost | Super-Apes | Mandarin | Bushmaster | U-Foes (Vector, Vapor, X-Ray & Ironclad) | Crossfire | Serpent Society | Rattler | Death Adder | Anaconda | Air-Walker

Avengers Assemble
2010 Marvel Animated Universe
Hydra/A.I.M. (Red Skull, M.O.D.O.K, Blood Brothers, Grim Reaper, Crossbones & Baron Wolfgang von Strucker) | J. Jonah Jameson | Doctor Doom | Ulik | Impossible Man | Dracula | Attuma | Wrecker | Thunderball | Piledriver | Bulldozer | Destroyer | Loki Laufeyson | Chitauri | Justin Hammer | Mojo | Galactus | Ringmaster | Black Order (Thanos) | Hela | Winter Soldier | Squadron Supreme (Nighthawk, Hyperion, Zarda, Speed Demon, Doctor Spectrum & Nuke) | Crimson Dynamo | Radioactive Man | Zzzax | Dormammu | Ultron | U-Foes (Vector, Vapor, X-Ray & Ironclad) | Roxxon Energy Corporation | Absorbing Man | Titania | Whirlwind | Fin Fang Foom | Maximus | Medusa | Masters of Evil/Thunderbolts (Helmut Zemo, Beetle, Screaming Mimi, Moonstone, Fixer & Goliath) | Klaw | Ghost | A.I.M. (Kang the Conqueror, Whiplash & Spymaster) | Abomination | Heinrich Zemo | Kree Empire | Egghead | The Leader | Crimson Widow | Arnim Zola | Enchantress | Skurge | Typhoid Mary | Ares | Taskmaster | Doughboy | Ulik | Beyonder | Baron Mordo | Symbiotes | Morgan le Fey

Eric Radomski Animated Universe
Shadow Council (Erik Killmonger, Helmut Zemo, Tiger Shark, Princess Zanda, M'Baku, Madame Masque, Klaw & Heinrich Zemo) | White Wolf | Hydra (Red Skull, Crossbones, Crimson Widow, Arnim Zola & Typhoid Mary) | Bask | Vulture | N'Basa | Kraven the Hunter | Taskmaster |Morgan le Fay | Orka | Winter Soldier | Attuma

Marvel Disk Wars: The Avengers
Loki Laufeyson | A.I.M. (M.O.D.O.K.) | Abomination | Baron Zemo | HYDRA (Red Skull, Crossbones) | King Cobra | Whirlwind | Tiger Shark | Whiplash | Diamondback | Crimson Dynamo | Dormammu | Sabretooth | Diablo | Graviton | Juggernaut | Lizard | Absoring Man | Wrecking Crew (Bulldozer, Piledriver, Thunderball, Wrecker) | Silver Samurai | Green Goblin (Norman Osborn) | Venom | Ultron | Cottonmouth | Doctor Octopus | Mandarin | Ronan | Fin Fang Foom | Baron Blood | Vampires | Bi-Beast | Obadiah Stane | Predator X | Mystique | Taskmaster | Blizzard

Marvel Future Avengers
Masters of Evil (Ares, Enchantress, Leader, Winter Soldier) | Loki Laufeyson | Kang the Conqueror | Maximus | HYDRA/A.I.M. (Red Skull, M.O.D.O.K., Arnim Zola) | Green Goblin (Norman Osborn) | Super-Adaptoid | Deadpool | Morgan le Fay | Klaw | Crossbones | Blizzard | Diamondback | The Hood | Ezekiel Stane

Marvel's M.O.D.O.K.
M.O.D.O.K. | A.I.M. | Monica Rappaccini | The Anomaly | Hexus | Super-Adaptoid | Austin Van Der Sleet | Mister Sinister | Madame Masque | Whirlwind | Mandrill | The Leader | Arcade | Fin Fang Foom | The Brood | Frost Giants

What If...?
Infinity Ultron | Yellowjacket | Loki Laufeyson | The Destroyer | Brock Rumlow | Jack Rollins | Thunderbolt Ross | Whiplash | Arnim Zola | Prince Killmonger | Strange Supreme | Georges Batroc | Ego | Zombies (Scarlet Witch)

Video Games
Captain America and The Avengers: Red Skull | Crossbones | Klaw | Living Laser | Whirlwind | Sentinel | Wizard | Grim Reaper | Mandarin | Juggernaut | Ultron
Avengers in Galactic Storm: Kree (Ronan, Korath & Supreme Intelligence)
Marvel Avengers: Battle for Earth: Skrulls (Queen Veranke & Super-Skrull) | Loki Laufeyson | Dark Phoenix | Venom (Eddie Brock) | Magneto | Ares | Norman Osborn
Marvel Disk Wars: The Avengers - Ultimate Heroes: Red Skull | Loki Laufeyson | Baron Zemo | A.I.M. (M.O.D.O.K.) | Abomination | Graviton | Tiger Shark | Green Goblin (Norman Osborn) | Crimson Dynamo | Whirlwind | Whiplash | Crossbones | Doctor Octopus | Serpent Society (King Cobra, Cottonmouth & Death Adder) | Wrecking Crew (Bulldozer, Piledriver, Thunderball & Wrecker) | Baron Mordo | Grey Gargoyle | Griffin | Fixer | Amora the Enchantress | Skurge the Exceutioner | Carnage | Klaw | Nightmare | Quicksilver | Scarlet Witch | Destroyer | Venom | Sandman | Winter Soldier | Super-Adaptoid | Taskmaster | Thanos | Rhino | Lizard | Absorbing Man | Deadpool
LEGO Marvel's Avengers: Loki Laufeyson | Malekith the Accursed | Kurse | Various Villains
Marvel's Avengers: M.O.D.O.K. | Monica Rappaccinni | Abomination | Taskmaster | Super-Adaptoid | Lyle Getz | Maestro | Klaw | Crossbones

See Also
Avengers Academy Villains | Marvel Avengers Alliance Villains | Young Avengers Villains

           Guardians of the Galaxy logo Villains

Comics
Abomination | Absorbing Man | A.I.M | Alkhema | Angela | Annihilation Wave | Annihilus | Apocalypse | Archangel | Badoon | Beyonder | Blastaar | Black Cat | Black Order | Blob | Blood Brothers | Brood | Bulldozer | Carnage | Collector | Celestials | Chitauri | Crime-Master | Dark Avengers | Dark Phoenix | Diablo | Doctor Doom | Doctor Octopus | Dormammu | Dragon of the Moon | Eddie Brock | Ego the Living Planet | Electro | Fin Fang Foom | Firelord | Frost Giants | Galactus | Garthan Saal | Gorgon | Grandmaster | Green Goblin | Grey Gargoyle | Hela | Hera | High Evolutionary | Jack O' Lantern | J'son | Juggernaut | Kang the Conqueror | Korath the Pursuer | Korvac | Knull | Kraven | Kree | Loki Laufeyson | Madame Masque | Maelstrom | Magneto | Magus | Mastermind | Masters of Evil | Maximus | Medusa | Mephisto | M.O.D.O.K. | Mole Man | Namor McKenzie | Nebula | Norman Osborn | Oddball | Piledriver | Puff Adder | Pyro | Raker | Red Skull | Ronan | Rhino | Sabretooth | Sandman | Savage Six | Sentinels | Shocker | Skrulls | Space Phantom | Super-Adaptoid | Super-Skrull | Supreme Intelligence | Symbiotes | Taskmaster | Terrax | Thanos | The Matriarch | Titania | Thunderball | Ultimus | Ultron | Universal Church of Truth | Venom | Vulcan | Vulture | Whiplash | Whirlwind | Wrecker | Xemnu

Movies
Guardians of the Galaxy: Kree Empire (Ronan the Accuser, Nebula, Korath the Pursuer, Sakaaran Mercenaries & Exolon Monks) | Yondu Ravager Clan (Yondu Udonta, Kraglin Obfonteri, Horuz & Vorker) | Garthan Saal | Eson the Searcher | Chitauri | Moloka Dar | Monstrous Inmate | The Collector | Dark Elf | Thanos
Guardians of the Galaxy Vol. 2: Ego | Sovereign (Ayesha, Sovereign Admiral, Zylak & Sovereign Chambermaid) | Yondu Ravager Clan (Yondu Udonta, Kraglin Obfonteri, Nebula, Tullk, Oblo, Taserface, Gef, Retch, Halfnut, Brahl, Vorker, Narblik & Huhtar) | Abilisks | The Grandmaster | Thanos
Guardians of the Galaxy Vol. 3: Orgocorp (High Evolutionary, Theel, Vim, Behemoth and War Pig, Controller, Karja, Kwol, Bletelsnort, Carrot Head, Ura, Phlektik, Orgosentries & Hell Spawn) | Sovereign (Ayesha & Adam Warlock) | Abilisks

TV Series
Guardians of the Galaxy (TV Series): Thanos | J'son | Kree | Ronan the Accuser | Korath the Pursuer | Nebula | Black Order | Loki Laufeyson | Destroyer | Medusa | Maximus | High Evolutionary | Supreme Intelligence | Symbiotes | Chitauri | Magus | Blood Brothers | Fin Fang Foom | The Collector | Grandmaster | A.I.M (M.O.D.O.K) | Venom | Carnage | Hela
What If...? (Season 1): Infinity Ultron | The Collector Ego the Living Planet (T'Challa Star-Lord)|Ego| Black Order (Ebony Maw, Proxima Midnight, Corvus Glaive & Cull Obsidian) | Ego | Prince Killmonger | Arnim Zola | Strange Supreme
What If...? (Season 2): Irani Rael | Ego | Yondu

Video Games
Marvel's Guardians of the Galaxy: Magus | Universal Church of Truth (Grand Unifier Raker, Nicholette Gold & Inquisitor) | Lady Hellbender | Thanos | Dweller-In-Darkness | Lethal Legion (Glah-Ree & Blood Brothers) | Wendigo | Fin Fang Foom | The Aliens (Solar Skies)

See Also
Howard the Duck Villains | Nova Villains

           Captain Marvel (2018) logo Villains

Comics
Absorbing Man | A.I.M. | Annihilus | Awesome Android | Badoon | Blastaar | Brood | Centurion | Controller | Dark Avengers | Deathbird | Doctor Faustus | Dormammu | Electro | Elementals | Enchantress | Frost Giants | Galactus | Grant Ward | Grotesk | Kang | King Cobra | Kree | Korath the Pursuer | Living Laser | Loki | Maestro | Molecule Man | Mandarin | Mercurio | Melter | Minn-Erva | M.O.D.O.K. | Moonstone | Mystique | Nebula | Nitro | Ove | Porcupine | Psycho-Man | Puppet Master | Quasimodo | Red Ghost | Rogue | Ronan | Scorpion | Sentry 459 | Skrulls | Skurge | Starforce | Star | Stranger | Super-Adaptoid | Super-Apes | Super-Skrull | Supreme Intelligence | Talos the Untamed | Thanos | Tiger Shark | The Omen (Leonore) | The Undone | Ultimus | Vampires | Yon-Rogg

Movies
Captain Marvel: Kree Empire/Starforce (Yon-Rogg, Minn-Erva, Korath the Pursuer, Att-lass, Bron-Char, Ronan the Accuser, Soh-Larr & Supreme Intelligence) | Skrulls (Talos & Norex) | Thanos
The Marvels: Kree Empire (Dar-Benn, Ty-Rone, Supreme Intelligence & Yon-Rogg)

See Also
Ms. Marvel Villains

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