Tyler is first seen delivering an Allinol tank to be attached to Rod Torque Redline. Then, he was smiling when Rod Torque Redline was going to be hit by the electric pulse camera.
Eventually, Finn McMissile came over to Lightning McQueen’s pit to stop Acer and Petey Pacer from entering and attacking Mater. Once Finn sprayed them with foam from a fire extinguisher and began to flee, countless members of the lemons started to chase him.
While Tyler was driving silently down the road looking for Mater, Finn used his hook to knock over a vending machine in front of Tyler, who took the blame for it.
Like all the other Lemons, Tyler Gremlin is a cold-hearted terrorist who would do anything for a quick buck. Apparently, he is very homicidal because he was involved in the death of Rod Torque Redline.
He is also very sadistic for attempting to abduct Mater until Hans and two other vehicles yelled at him.